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Question then, is the F2.8 worth the extra £300 over and above the F4
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Well it certainly would be to some people but that doesn't mean it is to you. After all £300 may not be all that much in terms of L glass but it's a big chunk of cash.
The picture quality of the two side by side, f stop for f stop is not going to be all that different. Much less than say the 300 f4L and the 300 f2.8L . Where you will most likely see more difference is in the focusing speed and how well it works with a converter on.
The f4 works pretty well with the 1.4 converter but will not AF with a (canon) 2x. The f2.8 is not going to have that issue and will always be brighter to frame with.
The other thing which could make a big difference is the weight. The f4 is compact and easy to live with compared to the 2.8